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The FX45 incorporates a 4.5-liter (4,494 cc) 32-valve V8 DOHC VK45DE gasoline engine with four valves per cylinder, microfinished camshafts with aluminum-alloy block and heads, low-friction molybdenum-coated pistons, titanium intake and exhaust valves, modular cylinder heads, microfinished crankshaft, lightweight pistons, super-silent single stage cam drive chain, and a Continuously Variable Valve Timing Control System that optimizes opening of intake valves.

The FX35 is available in two drivetrains, the RWD and the AWD, which are both fitted to a dual overhead cam 3.5-liter 24-valve V6 VQ35DE gasoline engine, with aluminum-alloy block and heads, electronically controlled “drive-by-wire” throttle system, and low-friction molybdenum-coated pistons that makes use of the continuous variable valve timing which in part optimizes the opening of intake valves. This is also coupled with an electronically controlled super-wide ratio five-speed automatic with a manumatic shift mode that includes the option of sequentially selected manual gearshifts and Downshift Rev Matching.

The truck is propelled by a 5.6 L VK56DE V8 engine (315 hp [227 kW] @ 4,900 rpm and 390 lb·ft (529 N·m) torque @ 3,600 rpm) and a 5-speed automatic transmission with an overdrive. Each cylinder in the engine is lined with cast iron. The valvetrain is a DOHC design with four valves per cylinder. Infiniti designed the QX56′s throttle responsiveness to correspond to specific drivetrain modes: when the driver selects all-wheel drive, the throttle becomes less sensitive, presumably to allow easier regulation of acceleration when off-road.